About the Authors

Heidi J Auman

Affiliation Developmental Genetics Program and Department of Cell Biology, Skirball Institute of Biomolecular Medicine, New York University School of Medicine, New York, New York, United States of America

Hope Coleman

Affiliation Developmental Genetics Program and Department of Cell Biology, Skirball Institute of Biomolecular Medicine, New York University School of Medicine, New York, New York, United States of America

Heather E Riley

Affiliation Developmental Genetics Program and Department of Cell Biology, Skirball Institute of Biomolecular Medicine, New York University School of Medicine, New York, New York, United States of America

Felix Olale

Affiliation Developmental Genetics Program and Department of Cell Biology, Skirball Institute of Biomolecular Medicine, New York University School of Medicine, New York, New York, United States of America

Huai-Jen Tsai

Affiliation Institute of Molecular and Cell Biology, College of Life Science, National Taiwan University, Taipei, Taiwan

Deborah Yelon

To whom correspondence should be addressed. E-mail: yelon@saturn.med.nyu.edu

Affiliation Developmental Genetics Program and Department of Cell Biology, Skirball Institute of Biomolecular Medicine, New York University School of Medicine, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

HJA, HC, and DY conceived and designed the experiments. HJA, HC, HER, and DY performed the experiments and analyzed the data. FO and HJT contributed reagents/materials/analysis tools. HJA and DY wrote the paper.