About the Authors

Sagiv Shifman, Jordana Tzenova Bell, Richard R Copley, Martin S Taylor, Richard Mott, Jonathan Flint
Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Robert W Williams
Department of Anatomy and Neurobiology, University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

Corresponding Author

jf@well.ox.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

RWW, RM, and JF conceived and designed the experiments. SS, JTB RRC, MST, and RM analyzed the data. RWW and RM contributed reagents/materials/analysis tools. SS, JTB, and JF wrote the paper. SS and JTB performed data cleaning and statistical analysis. RRC and MST performed the sequence analysis. RW coordinated sample collection. RM and JF formulated and directed the project.