About the Authors

Matthew J Marshall, Alexander S Beliaev, Alice C Dohnalkova, David W Kennedy, Liang Shi, Jeffrey S McLean, Samantha B Reed, David E Culley, Vanessa L Bailey, Cody J Simonson, Margaret F Romine, James K Fredrickson
Biological Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Zheming Wang, John M Zachara
Chemical Sciences Division, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Maxim I Boyanov, Kenneth M Kemner
Biosciences Division, Argonne National Laboratory, Argonne, Illinois, United States of America

Barry Lai
Advanced Photon Source, Argonne National Laboratory, Argonne, Illinois, United States of America

Daad A Saffarini
Department of Biological Sciences, University of Wisconsin, Milwaukee, Wisconsin, United States of America

Corresponding Authors

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MJM, ASB, KMK, JKF, and JMZ conceived and designed the experiments. MJM, ACD, DWK, LS, ZW, MIB, BL, KMK, JSM, SBR, DEC, and CJS performed the experiments. MJM, ASB, KMK, VLB, JKF, and JMZ analyzed the data. DAS and MFR contributed reagents/materials/analysis tools. MJM, ASB, JKF, and JMZ wrote the paper.