About the Authors

Ilkka Hanski

To whom correspondence should be addressed. E-mail: ilkka.hanski@helsinki.fi

Affiliation Department of Biological and Environmental Sciences, University of Helsinki, Finland

Ilik Saccheri

Affiliation School of Biological Sciences, University of Liverpool, Liverpool, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

IH collected the material. IS genotyped the larval material. IH analysed the data. IH and IS wrote the paper.