About the Authors

Alexei J Drummond, Simon Y. W Ho, Matthew J Phillips, Andrew Rambaut
Department of Zoology, University of Oxford, Oxford, United Kingdom

Corresponding Author

andrew.rambaut@zoo.ox.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AJD and AR conceived the original idea, developed the software, and performed the marsupial and virus data analyses. SYWH developed the simulation software, performed the simulation analysis, developed the use of prior distributions for calibrating node ages, and performed the analyses on the bacteria, yeast, plant, metazoan, and primate datasets. MJP collected and curated the marsupial dataset and provided expert calibration information. AJD, SYWH, MJP, and AR contributed to the writing of the article.