About the Authors

Min Hu

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Lichuan Gu

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Muyang Li

Affiliation Institute for Cancer Genetics, Department of Pathology, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Philip D Jeffrey

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Wei Gu

Affiliation Institute for Cancer Genetics, Department of Pathology, Columbia University College of Physicians and Surgeons, New York, New York, United States of America

Yigong Shi

To whom correspondence should be addressed. E-mail: yshi@molbio.princeton.edu

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MH and YS conceived and designed the experiments. MH, LG, and PDJ performed the experiments. MH, LG, PDJ, and YS analyzed the data. MH, LG, ML, PDJ, and WG contributed reagents/materials/analysis tools. MH and YS wrote the paper.