About the Authors

Douglas M Fowler

Contributed equally to this work with: Douglas M Fowler, Atanas V Koulov

Affiliation Department of Chemistry and The Skaggs Institute of Chemical Biology, The Scripps Research Institute, La Jolla, California, United States of America

Atanas V Koulov

Contributed equally to this work with: Douglas M Fowler, Atanas V Koulov

Affiliation Department of Cell Biology and the Institute for Childhood and Neglected Diseases, The Scripps Research Institute, La Jolla, California, United States of America

Christelle Alory-Jost

Affiliation Department of Cell Biology and the Institute for Childhood and Neglected Diseases, The Scripps Research Institute, La Jolla, California, United States of America

Michael S Marks

Affiliation Department of Pathology and Laboratory Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

William E Balch

Affiliation Department of Cell Biology and the Institute for Childhood and Neglected Diseases, The Scripps Research Institute, La Jolla, California, United States of America

Jeffery W Kelly

To whom correspondence should be addressed. E-mail: jkelly@scripps.edu

Affiliation Department of Chemistry and The Skaggs Institute of Chemical Biology, The Scripps Research Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

WEB and JWK conceived and designed the experiments. DMF, AVK, and CA performed the experiments. DMF, AVK, and CA analyzed the data. MSM, WEB, and JWK wrote the paper.