About the Authors

Donald C Cooper

Affiliations Department of Neurobiology and Physiology, Institute for Neuroscience, Northwestern University, Evanston, Illinois, United States of America, Department of Psychiatry, University of Texas Southwestern Medical, Dallas, Texas, United States of America

Sungkwon Chung

Affiliations Department of Neurobiology and Physiology, Institute for Neuroscience, Northwestern University, Evanston, Illinois, United States of America, Department of Physiology, Sungkyunkwan University School of Medicine, Suwan, South Korea

Nelson Spruston

To whom correspondence should be addressed. E-mail: spruston@northwestern.edu

Affiliation Department of Neurobiology and Physiology, Institute for Neuroscience, Northwestern University, Evanston, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DCC and NS conceived and designed the experiments. DCC and SC performed the experiments. DCC analyzed the data and wrote the paper with NS.