About the Authors

Fiona Oliver, Julian K Christians, Xiaojun Liu, Vinesh Verma, Peter D Keightley
University of Edinburgh, Institute of Evolutionary Biology, School of Biological Sciences, Edinburgh, United Kingdom

Susan Rhind
Royal (Dick) School of Veterinary Studies, Easter Bush Veterinary Centre, Roslin, Midlothian, United Kingdom

Claire Davison, Steve D. M Brown, Paul Denny
Medical Research Council Mammalian Genetics Unit, Harwell, Oxfordshire, United Kingdom

Corresponding Author

julian.christians@ed.ac.uk

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

FO, JKC, SDMB, PD, and PDK conceived and designed the experiments. FO, JKC, XL, SR, VV, and CD performed the experiments. FO, JKC, and PDK analyzed the data. FO, JKC, and PDK wrote the paper.