About the Authors

Graham E. J Rodwell

Affiliation Division of Nephrology, Stanford University Medical Center, Stanford, California, United States of America

Rebecca Sonu

Affiliation Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America

Jacob M Zahn

Affiliation Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America

James Lund

Affiliation Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America

Julie Wilhelmy

Affiliation Department of Biochemistry, Stanford University Medical Center, Stanford, California, United States of America

Lingli Wang

Affiliation Department of Pathology, Stanford University Medical Center, Stanford, California, United States of America

Wenzhong Xiao

Affiliation Department of Biochemistry, Stanford University Medical Center, Stanford, California, United States of America

Michael Mindrinos

Affiliation Department of Biochemistry, Stanford University Medical Center, Stanford, California, United States of America

Emily Crane

Affiliation Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America

Eran Segal

Affiliation Department of Computer Science, Stanford University Medical Center, Stanford, California, United States of America

Bryan D Myers

Affiliation Division of Nephrology, Stanford University Medical Center, Stanford, California, United States of America

James D Brooks

Affiliation Department of Urology, Stanford University Medical Center, Stanford, California, United States of America

Ronald W Davis

Affiliations Department of Biochemistry, Stanford University Medical Center, Stanford, California, United States of America, Department of Genetics, Stanford University Medical Center, Stanford, California, United States of America

John Higgins

Affiliation Department of Pathology, Stanford University Medical Center, Stanford, California, United States of America

Art B Owen

Affiliation Department of Statistics, Stanford University Medical Center, Stanford, California, United States of America

Stuart K Kim

To whom correspondence should be addressed. E-mail: kim@cmgm.stanford.edu

Affiliations Department of Developmental Biology, Stanford University Medical Center, Stanford, California, United States of America, Department of Genetics, Stanford University Medical Center, Stanford, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

SKK conceived and designed the experiments. RS, JW, LW, EC, and JH performed the experiments. SKK, GEJR, JMZ, JL, WX, MM, ES, and ABO analyzed the data. MM, BDM, JDB, RWD and JH contributed reagents/materials/analysis tools. SKK, JMZ, and ABO wrote the paper.