About the Authors

Andrew Fortna

Contributed equally to this work with: Andrew Fortna, Young Kim

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Young Kim

Contributed equally to this work with: Andrew Fortna, Young Kim

Affiliation Department of Pathology, Stanford University, Stanford, California, United States of America

Erik MacLaren

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Kriste Marshall

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Gretchen Hahn

Affiliation Colorado Genetics Laboratory, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Lynne Meltesen

Affiliation Colorado Genetics Laboratory, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Matthew Brenton

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Raquel Hink

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Sonya Burgers

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Tina Hernandez-Boussard

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Anis Karimpour-Fard

Affiliation Department of Preventive Medicine and Biometrics, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Deborah Glueck

Affiliation Department of Preventive Medicine and Biometrics, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Loris McGavran

Affiliation Colorado Genetics Laboratory, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Rebecca Berry

Affiliation Colorado Genetics Laboratory, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Jonathan Pollack

To whom correspondence should be addressed. E-mail: pollack1@stanford.edu (JP), Email: james.sikela@uchsc.edu (JS)

Affiliation Department of Pathology, Stanford University, Stanford, California, United States of America

James M Sikela

To whom correspondence should be addressed. E-mail: pollack1@stanford.edu (JP), Email: james.sikela@uchsc.edu (JS)

Affiliation Department of Pharmacology and Human Medical Genetics Program, University of Colorado Health Sciences Center, Denver, Colorado, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

JP and JS conceived and designed the experiments. AF, YK, EM, and GH performed the experiments. AF, EM, KM, LM, MB, SB, AK-F, DG, LM, RB, JP, and JS analyzed the data. AF, YK, EM, RH, TH-B, and JP contributed reagents/materials/analysis tools. JS wrote the paper.