About the Authors

Pamela F Colosimo

Contributed equally to this work with: Pamela F Colosimo, Catherine L Peichel

Affiliation Department of Developmental Biology and Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Catherine L Peichel

Contributed equally to this work with: Pamela F Colosimo, Catherine L Peichel

Affiliation Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Kirsten Nereng

Affiliation Department of Developmental Biology and Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Benjamin K Blackman

Affiliation Department of Developmental Biology and Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Michael D Shapiro

Affiliation Department of Developmental Biology and Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Dolph Schluter

Affiliation University of British Columbia, Vancouver, British Columbia, Canada

David M Kingsley

To whom correspondence should be addressed. E-mail: kingsley@cmgm.stanford.edu

Affiliation Department of Developmental Biology and Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

PFC, CLP, DS, and DMK conceived and designed the experiments. PFC, CLP, KN, BJB, MDS, and DS performed the experiments. PFC, CLP, and DMK analyzed the data. DS contributed reagents/materials/analysis tools. PFC, CLP, MDS, and DMK wrote the paper.