About the Authors

Howard Y Chang

Affiliations Department of Dermatology, Stanford University School of Medicine, Stanford, California, United States of America, Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Julie B Sneddon

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Ash A Alizadeh

Current address: Department of Microbiology and Immunology, University of California at San Francisco, San Francisco, California, United States of America

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Ruchira Sood

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Rob B West

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Kelli Montgomery

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Jen-Tsan Chi

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Matt van de Rijn

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

David Botstein

Current address: Lewis–Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Patrick O Brown

To whom correspondence should be addressed. E-mail: pbrown@pmgm2.stanford.edu

Affiliations Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

HYC, DB, and POB conceived and designed the experiments. HYC, JBS, RS, and KM performed the experiments. HYC, JBS, AAA, RBW, JTC, MVDR, DB, and POB analyzed the data. HYC and POB wrote the paper.