About the Authors

Jeroen P Roose

Affiliations Department of Medicine, University of California, San Francisco, San Francisco, California, United States of America, Department of Microbiology and Immunology, University of California, San Francisco, San Francisco, California, United States of America

Maximilian Diehn

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Michael G Tomlinson

Affiliations Department of Medicine, University of California, San Francisco, San Francisco, California, United States of America, Department of Microbiology and Immunology, University of California, San Francisco, San Francisco, California, United States of America

Joseph Lin

Affiliations Department of Medicine, University of California, San Francisco, San Francisco, California, United States of America, Department of Microbiology and Immunology, University of California, San Francisco, San Francisco, California, United States of America

Ash A Alizadeh

Address address: Department of Microbiology and Immunology, University of California, San Francisco, San Francisco, California, United States of America

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

David Botstein

Address address: Lewis-Sigler Institute, Princeton University, Princeton, New Jersey, United States of America

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Patrick O Brown

Affiliations Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Arthur Weiss

To whom correspondence should be addressed. E-mail: aweiss@medicine.ucsf.edu

Affiliations Department of Medicine, University of California, San Francisco, San Francisco, California, United States of America, Department of Microbiology and Immunology, University of California, San Francisco, San Francisco, California, United States of America, Howard Hughes Medical Institute, University of California, San Francisco, San Francisco, California, United States of America, Rosalind Russell Medical Research Center for Arthritis, University of California, San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

JPR and AW conceived and designed the experiments. JPR, MD, MGT, JL, and AAA performed the experiments. JPR, MD, MGT, and AAA analyzed the data. MD, JL, DB, and POB contributed reagents/materials/analysis tools. JPR wrote the paper. POB assisted in preparing the manuscript. AW guided JPR's project and assisted in writing the manuscript.