About the Authors

Hong-Ling Huang

Contributed equally to this work with: Hong-Ling Huang, Shimin Wang

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Shimin Wang

Contributed equally to this work with: Hong-Ling Huang, Shimin Wang

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Meng-Xin Yin

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Liang Dong

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Chao Wang

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Wei Wu

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Yi Lu

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Miao Feng

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Chuanyang Dai

Affiliation Shanghai Ocean University, Shanghai, P.R. China

Xiaocan Guo

Affiliation Life Sciences Institute, Zhejiang University, Hangzhou, Zhejiang, P.R. China

Li Li

Affiliation Institute of Ageing Research, Hangzhou Normal University, Hangzhou, Zhejiang, P.R. China

Bin Zhao

Affiliation Life Sciences Institute, Zhejiang University, Hangzhou, Zhejiang, P.R. China

Zhaocai Zhou

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Hongbin Ji

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Jin Jiang

Affiliation Department of Developmental Biology, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America

Yun Zhao

yunzhao@sibcb.ac.cn (YZ); xyliu@sibcb.ac.cn (XYL); rayzhang@sibcb.ac.cn (LZ)

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Xin-Yuan Liu

yunzhao@sibcb.ac.cn (YZ); xyliu@sibcb.ac.cn (XYL); rayzhang@sibcb.ac.cn (LZ)

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Lei Zhang

yunzhao@sibcb.ac.cn (YZ); xyliu@sibcb.ac.cn (XYL); rayzhang@sibcb.ac.cn (LZ)

Affiliation State Key Laboratory of Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, Shanghai, P.R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: HLH SW XYL LZ. Performed the experiments: HLH SW LD XG. Analyzed the data: HLH SW MXY BZ ZZ HJ JJ YZ XYL LZ. Contributed reagents/materials/analysis tools: CW WW YL MF CD LL BZ ZZ HJ JJ YZ. Wrote the paper: HLH SW MXY LZ.