About the Authors

Stephen J Huston

Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom , Department of Bioengineering, Imperial College London, London, United Kingdom , Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Holger G Krapp

To whom correspondence should be addressed. E-mail: h.g.krapp@imperial.ac.uk

Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom , Department of Bioengineering, Imperial College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

SJH and HGK conceived and designed the experiments. SJH performed the experiments and analyzed the data. SJH and HGK wrote the paper.