About the Authors

Youbin Xiang

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Satomi Takeo

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Laurence Florens

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Stacie E Hughes

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Li-Jun Huo

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

William D Gilliland

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Selene K Swanson

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Kathy Teeter

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Joel W Schwartz

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Michael P Washburn

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Sue L Jaspersen

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America , University of Kansas School of Medicine, Kansas City, Kansas, United States of America

R. Scott Hawley

To whom correspondence should be addressed. E-mail: RSH@stowers-institute.org

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America , University of Kansas School of Medicine, Kansas City, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

YX, ST, LF, SEH, LJH, WDG, SKS, JWS, MPW, SLJ and RSH conceived and designed the experiments. YX, ST, LF, SEH, LJH, WDG, SKS, KT, JWS, MPW, and SLJ performed the experiments. LF analyzed the data. RSH wrote the manuscript in collaboration with all other authors.