About the Authors

Marc Hammarlund

Contributed equally to this work with: Marc Hammarlund, Mark T Palfreyman, Shigeki Watanabe

Affiliations Department of Biology, University of Utah, Salt Lake City, Utah, United States of America , Howard Hughes Medical Institute, University of Utah, Salt Lake City, Utah, United States of America

Mark T Palfreyman

Contributed equally to this work with: Marc Hammarlund, Mark T Palfreyman, Shigeki Watanabe

Affiliations Department of Biology, University of Utah, Salt Lake City, Utah, United States of America , Howard Hughes Medical Institute, University of Utah, Salt Lake City, Utah, United States of America

Shigeki Watanabe

Contributed equally to this work with: Marc Hammarlund, Mark T Palfreyman, Shigeki Watanabe

Affiliations Department of Biology, University of Utah, Salt Lake City, Utah, United States of America , Howard Hughes Medical Institute, University of Utah, Salt Lake City, Utah, United States of America

Shawn Olsen

Affiliations Department of Biology, University of Utah, Salt Lake City, Utah, United States of America , Howard Hughes Medical Institute, University of Utah, Salt Lake City, Utah, United States of America

Erik M Jorgensen

To whom correspondence should be addressed. E-mail: jorgensen@biology.utah.edu

Affiliations Department of Biology, University of Utah, Salt Lake City, Utah, United States of America , Howard Hughes Medical Institute, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MH, MTP, and EMJ conceived and designed the experiments and wrote the paper. MH, MTP, SW, and SO performed the experiments. MH, MTP, SW, and EMJ analyzed the data.