About the Authors

Joshua M Akey

To whom correspondence should be addressed. E-mail: jakey@fhcrc.org (JA), Email: leonid@fhcrc.org (LK)

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Michael A Eberle

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Mark J Rieder

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Christopher S Carlson

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Mark D Shriver

Affiliation Department of Anthropology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Deborah A Nickerson

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Leonid Kruglyak

To whom correspondence should be addressed. E-mail: jakey@fhcrc.org (JA), Email: leonid@fhcrc.org (LK)

Affiliations Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

MJR, CSC, and DAN conceived and designed the experiments. JMA, MAE, MDS, and LK analyzed the data. JMA, DAN, and LK wrote the paper.